The complex nature of superimposed collisional imprints across the Carpathian-Balkans and Dinaride-Hellenide mainly directs the research focus toward the late Alpine tectonically driven exhumation events (Neotethyan closure and subsequent Adria-Europe collision). So far, efforts to distinguish the tectonic stages before the Alpine event, which caused the displacement of older medium- to high-grade metamorphic basement rocks, have provided limited explanations. The main challenge lies in the transition between Variscan and pre-Variscan structural imprints. The exposed cluster of metamorphosed Cadomian-derived Cenerian and Variscan basement inliers, including their paleogeography, the timeline of deformation events, and the gradual development of deformation, remains poorly understood. To address these issues, we invite contributions from across the Carpathians as a whole, as well as from the Balkan Peninsula, including the Aegean/East Mediterranean/North African/Caucasus/Iranian north Gondwanan fragments, to help improve our understanding of the paleogeographic and geodynamic processes related to the evolution of these Paleozoic oceanic segments and their microcontinents. We welcome integrated petrological, structural, geochronological, and geochemical regional studies, as well as subsurface modeling efforts involving the overprinted Paleozoic microcontinents and vanished paleo-oceans, involving the Rheic, Galitia-Moldanubian (plus Hanseatic), and Variscan Paleotethys (including the Eocimmerian event).